Hayao Miyazaki's film, Princess Mononoke is about war, industrialization, and nature; it centers around two people, San and Ashitaka. San is a "wolf girl" who was raised by Moro, an ancient forest god, and Ashitaka comes from a peaceful people that's nearly gone. They both represent a dying world, but San sees Ashitaka as opposition because he's staying in Irontown. It's their alliance, that helps bring an end to a great curse, representing union and a symbiotic relationship. Studio Ghibli doesn't make many outright romance films, but the romances the studio produces are often subtle, beautiful, and say something about the best parts of humanity.
Source：https://www.cbr.com/princess-mononoke-perfect-romantic-subtext/
